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Bothremydidae (Redirected from Bothremydid)

Bothremydidae
Temporal range: Aptian–Miocene
Gafsachelys.jpg
Gafsachelys spp. skull
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Order: Testudines
Suborder: Pleurodira
Hyperfamily: Pelomedusoides
Family: Bothremydidae
Baur, 1891
Subfamilies

See text

Bothremydidae is an extinct family of side-necked turtles (Pleurodira) known from the Cretaceous and Cenozoic. They are closely related to Podocnemididae, and are amongst the most widely distributed pleurodire groups, with their fossils having been found in Africa, India, the Middle East, Europe, North America and South America. Bothremydids were aquatic turtles with a high morphological diversity, indicative of generalist, molluscivorous, and piscivorous diets. Unlike modern pleurodires, which are exclusively freshwater, bothremydids inhabited freshwater, marine and coastal settings. Their marine habits allowed bothremydids to disperse across oceanic barriers into Europe and North America during the early Late Cretaceous (Cenomanian). The youngest records of the group are indeterminate remains from Saudi Arabia and Oman, dating to the Miocene.

Taxonomy

The family is split into two subfamilies and a number of tribes.

Bothremydidae

  • Pleurochayah appalachius Adrian et al., 2021 Woodbine Formation, Texas, Late Cretaceous (Cenomanian)
  • "Chrysemys" montolivensis Roman 1897 Montoulieu, France, Oligocene (Rupelian)
  • Sindhochelys ragei Lapparent de broin et al., 2021 Khadro Formation, Pakistan, Paleocene (Danian)
  • Subfamily Bothremydinae Gaffney et al., 2006
    • Tribe Bothremydini Gaffney et al., 2006
    • Palauchelys López-Conde et al. 2021 Olmos Formation, Mexico, Late Cretaceous (Campanian)
      • Subtribe Bothremydina Gaffney et al., 2006
      • Subtribe Foxemydina Gaffney et al., 2006
      • Tribe Taphrosphyini Gaffney et al., 2006
          • Ilatardia Pérez-García, 2019 Farin Doutchi Formation, Niger, Late Cretaceous (Maastrichtian)
        • Subtribe Nigermydina Gaffney et al., 2006
          • Arenila Lapparent de Broin and Werner, 1998 Dakhla Formation, Egypt, Late Cretaceous (Maastrichtian)
            • Arenila krebsi Lapparent de Broin and Werner, 1998
        • Subtribe Taphrosphyina Gaffney et al., 2006
          • Azabbaremys Gaffney et al., 2001 Teberemt Formation, Mali, Paleocene
            • Azabbaremys moragjonesi Gaffney et al., 2001
          • Eotaphrosphys Perez-Garcia, 2018 Mont-Aimé, France, Late Cretaceous (Maastrichtian)
            • Eotaphrosphys ambiguum (Gaudry, 1890)
          • Labrostochelys Gaffney et al., 2006 Ouled Abdoun Basin, Morocco, Paleocene
            • Labrostochelys galkini Gaffney et al., 2006
          • Motelomama Perez-Garcia, 2018 Salina Group, Peru, Eocene (Ypresian)
            • Motelomama olssoni (Schmidt, 1931)
          • Phosphatochelys Gaffney and Tong, 2003 Ouled Abdoun Basin, Morocco, Paleocene
            • Phosphatochelys tedfordi Gaffney and Tong, 2003
          • Rhothonemys Gaffney et al., 2006 Ouled Abdoun Basin, Morocco, Paleocene
            • Rhothonemys brinkmani Gaffney et al., 2006
          • Taphrosphys Cope, 1869 (jr synonyms: Amblypeza, Bantuchelys, Prochonias)
            • Taphrosphys congolensis (Dollo, 1912) Landana Formation, Angola, Paleocene
            • Taphrosphys dares Hay, 1908 (nomen dubium)
            • Taphrosphys ippolitoi Gaffney et al., 2006 Ouled Abdoun Basin, Morocco, Paleocene
            • Taphrosphys sulcatus (Leidy, 1856) Hornerstown Formation, New Egypt Formation, New Jersey, Late Cretaceous (Maastrichtian)
          • Ummulisani Gaffney et al., 2006 Morocco, Eocene (Ypresian)
            • Ummulisani rutgersensis Gaffney et al., 2006
          • Crassachelys Bergounioux, 1952 (nomen dubium)
            • Crassachelys neurirregularis Bergounioux, 1952
          • Eusarkia Bergounioux, 1952 (nomen dubium)
            • Eusarkia rotundiformis Bergounioux, 1952
          • Gafsachelys Stefano, 1903 (nomen dubium)
            • Gafsachelys phosphatica Stefano, 1903
            • Gafsachelys moularensis Bergounioux, 1955
    • Tribe Cearachelyini Gaffney et al., 2006
      • Cearachelys Gaffney et al., Santana Formation, Brazil, Early Cretaceous (Albian)
        • Cearachelys placidoi Gaffney et al., 2001
      • Elkanemys Maniel, de la Fuente and Canale, 2021 Candeleros Formation, Argentina, Late Cretaceous (Cenomanian)
        • Elkanemys pritchardi Maniel, de la Fuente and Canale, 2021
      • Itapecuruemys Batista et al., 2020 Itapecuru Formation, Brasil, Early Cretaceous (Aptian)
        • Itapecuruemys amazoniensis Batista et al., 2020
      • Galianemys Gaffney et al., 2002 Kem Kem Group, Morocco, Late Cretaceous (Cenomanian)
        • Galianemys emringeri Gaffney et al., 2002
        • Galianemys whitei Gaffney et al., 2002
  • Subfamily Kurmademydinae Gaffney et al., 2006
    • Tribe Kurmademydini Gaffney et al., 2006

Phylogeny

Below is a cladogram by Gaffney et al. in 2006:

Chelidae

Pelomedusoides

Araripemys

Pelomedusidae

Euraxemydidae

Dirqadium

Euraxemys

Podocnemidioidea

Podocnemidinura

Bothremydidae
Kurmademydini

Kurmademys

Sankuchemys

Bothremydinae
Cearachelyini

Galianemys

Cearachelys

Bothremyodda
Bothremydini

Foxemys (=Polysternon)

Polysternon

Zolhafah

Rosasia

Araiochelys

Chedighaii hutchisoni

Chedighaii barberi

Bothremys cooki

Bothremys magrhebiana

Bothremys kellyi

Bothremys arabicus

Taphrosphyini

Nigeremys

Arenila

Azabbaremys

Labrostochelys

Taphrosphys

Rhothonemys

Phosphatochelys

Ummulisani


This page was last updated at 2022-01-14 06:59 UTC. Update now. View original page.

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